Botanic
fruits and culinary fruits
Many foods are botanically fruits. These
include cucurbits (e.g., squash and
pumpkin),
tomato, cucumber, aubergine (eggplant),
and sweet pepper, along with nuts, and some
spices, such as allspice, nutmeg and chiles.
Rarely, culinary "fruits" are not fruits in
the botanical sense. For example, rhubarb
may be considered a fruit, though only the
astringent stalk or petiole is edible. In
the commercial world, European Union rules
define carrot as a fruit for the purposes of
measuring the proportion of "fruit"
contained in carrot jam.
Fruit development
After an ovule is fertilized in a process
known as pollination, the ovary begins to
expand. The petals of the flower fall off
and the ovule develops into a seed. The
ovary eventually comes to form, along with
other parts of the flower in many cases, a
structure surrounding the seed or seeds that
is the fruit. Fruit development continues
until the seeds have matured. With some
multiseeded fruits the extent of development
of the flesh of the fruit is proportional to
the number of fertilized ovules.
The wall of the fruit, developed from the
ovary wall of the flower, is called the
pericarp. The pericarp is often
differentiated into two or three distinct
layers called the exocarp (outer layer -
also called epicarp), mesocarp (middle
layer), and endocarp (inner layer). In some
fruits, especially simple fruits derived
from an inferior ovary, other parts of the
flower (such as the floral tube, including
the petals, sepals, and stamens), fuse with
the ovary and ripen with it. When such other
floral parts are a significant part of the
fruit, it is called an accessory fruit.
Since other parts of the flower may
contribute to the structure of the fruit, it
is important to study flower structure to
understand how a particular fruit forms.
Fruits are so varied in form and
development, that it is difficult to devise
a classification scheme that includes all
known fruits. It will also be seen that many
common terms for seeds and fruit are
incorrectly applied, a fact that complicates
understanding of the terminology. Seeds are
ripened ovules; fruits are the ripened
ovularies or carpels that contain the seeds.
To these two basic definitions can be added
the clarification that in botanical
terminology, a nut is a type of fruit and
not another term for seed.
There are three basic types of fruits:
1. Simple
fruit
2. Aggregate fruit
3. Multiple fruit
Simple fruit
Simple fruits can be either dry or fleshy
and result from the ripening of a simple or
compound ovary with only one pistil. Dry
fruits may be either dehiscent (opening to
discharge seeds), or indehiscent (not
opening to discharge seeds). Types of dry,
simple fruits (with examples) are:
- achene -
(buttercup)
- capsule
- (Brazil nut)
-
caryopsis - (wheat)
- fibrous
drupe - (coconut, walnut)
- follicle
- (milkweed)
- legume -
(pea, bean, peanut)
- loment
- nut -
(hazelnut, beech, oak acorn)
- samara -
(elm, ash, maple key)
-
schizocarp - (carrot)
- silique
- (radish)
- utricle
Fruits in
which part or all of the pericarp (fruit
wall) is fleshy at maturity are simple
fleshy fruits. Types of fleshy, simple
fruits (with examples) are:
- berry -
(tomato,
avocado)
- Stone
fruit drupe (plum, cherry, peach, olive)
- false
berry - accessory fruits (banana,
cranberry)
- pome -
accessory fruits (apple, pear, rosehip)
Aggregate
fruit
An aggregate fruit, or etaerio, develops
from a flower with numerous simple pistils.
An example is the raspberry, whose simple
fruits are termed drupelets because each is
like a small drupe attached to the
receptacle. In some bramble fruits (such as
blackberry) the receptacle is elongate and
part of the ripe fruit, making the
blackberry an aggregate-accessory fruit. The
strawberry is also an aggregate-accessory
fruit, only one in which the seeds are
contained in achenes. In all these examples,
the fruit develops from a single flower with
numerous pistils.
Multiple fruit
A multiple fruit is one formed from a
cluster of flowers (called an
inflorescence). Each flower produces a
fruit, but these mature into a single mass.
Examples are the pineapple, edible fig,
mulberry, osage-orange, and breadfruit.
In
the photograph on the right, stages of
flowering and fruit development in the noni
or Indian mulberry (Morinda citrifolia) can
be observed on a single branch. First an
inflorescence of white flowers called a head
is produced. After fertilization, each
flower develops into a drupe, and as the
drupes expand, they connate (merge) into a
multiple fleshy fruit called a syncarp.
Seedless Fruits
Seedlessness is an important feature of some
fruits of commerce. Commercial cultivars of
bananas and pineapples are seedless. Some
cultivars of citrus fruits (especially navel
oranges and mandarin oranges), table grapes,
grapefruit, and watermelons are valued for
their seedlessness. In some species,
seedlessness is the result of parthenocarpy,
where fruits set without fertilization.
Parthenocarpic fruit set may or may not
require pollination. Most seedless citrus
fruits require a pollination stimulus;
bananas and pineapples do not. Seedlessness
in table grapes results from the abortion of
the embryonic plant that is produced by
fertilization, a phenomenon known as
stenospermocarpy which requires normal
pollination and fertilization.
Seed dissemination
Variations in fruit structures largely
relate to the mode of dispersal of the seeds
they contain.
Some fruits have coats covered with spikes
or hooked burrs, either to prevent
themselves from being eaten by animals or to
stick to the hairs of animals, using them as
dispersal agents. Other fruits are elongated
and flattened out naturally and so become
thin, like wings or helicopter blades. This
is an evolutionary mechanism to increase
dispersal distance away from the parent.
The sweet flesh of many fruits is
"deliberately" appealing to animals, so that
the seeds held within are "unwittingly"
carried away and deposited at a distance
from the parent. Likewise, the nutritious,
oily kernels of nuts are appealing to
rodents (such as squirrels) who hoard them
in the soil in order to avoid starving
during the winter, thus giving those seeds
that remain uneaten the chance to germinate
and grow into a new plant away from their
parent.
Uses
Many fruits, including fleshy fruits like
apples and mangos, and nuts like walnut, are
commercially valuable as human food, eaten
both fresh and made into jams, marmalade and
other preserves for future consumption.
Fruits are also found commonly in such
manufactured foods as cookies, muffins,
yoghurt, ice cream, cakes, and many more.
Mediterranean and subtropical fruits
Fruits in this category are not hardy to
extreme cold, as the preceding temperate
fruits are, yet tolerate some frost and may
have a modest chilling requirement. Notable
among these are natives of the
Mediterranean:
Cornelian cherry (Cornus mas; Cornaceae)
-
Date palm
(Phoenix dactylifera; Arecaceae)
-
Fig (Ficus spp.
Moraceae)
-
Grape, called
raisin, sultana, or currant when it is dried
(Vitis spp.; Vitaceae)
-
Jujube (Ziziphus
zizyphus; Rhamnaceae)
-
Black mulberry
(Morus nigra; Moraceae)
-
Olive (Olea
europea; Oleaceae)
-
Pomegranate (Punica
granatum; Punicaceae)
In the
important genus Citrus (Rutaceae), some
members are tropical, tolerating no frost.
All common species of commerce are somewhat
hardy:
-
Pomelo (also
known as the shaddock) (Citrus paradisi)
-
Citron (Citrus medica)
-
Grapefruit and
its predecesor the
-
Lemon (Citrus
limon)
- Limes
- Key
Lime (Citrus aurantifolia)
-
persian lime Also known as thaiti lime.
- kaffir
lime (Citrus hystix)
- Mandarin
(Citrus reticulata),
-
Key Lime
(Citrus aurantifolia)
-
persian lime
Also known as thaiti lime.
-
kaffir lime
(Citrus hystix)
-
Mandarin
(Citrus reticulata),
-
Clementine
(Citrus reticulata var. Clementine),
-
Tangerine, and
similar
-
Orange, of
which there are sweet (Citrus sinensis) and
sour (Citrus aurantium) species
-
Hybrids of the
preceding species, such as the Orangelo,
Tangelo, rangpur and Ugli fruit
Other
subtropical fruits:
-
Avocado (Persea
americana; Lauraceae)
-
Feijoa (Feijoa
sellowiana; Myrtaceae)
-
Guava (Psidium
guajava; Myrtaceae)
-
Kumquat (Fortunella
spp.; Rutaceae)
-
Longan
(Euphoria longan; Sapindaceae, the same
family as maples and buckeyes)
-
Lychee (Litchi
chinensis; Sapindaceae)
-
Passion fruit
or Grenadilla (Passiflora edulis and other
Passiflora spp.; Passifloraceae)
-
Strawberry
guava (Psidium litorale; Myrtaceae)
-
Tamarillo or
Tree Tomato (Cyphomandra betacea; Solanaceae)
Tropical
fruits
Tropical fruit grow on plants of all
habitats. The only characteristic that they
share is an intolerance of frost.
-
Acai (Euterpe
oleracea; Arecaceae)
-
African cherry
orange (Citropsis schweinfurthii; Rutaceae)
-
Akee (Blighia
sapida or Cupania sapida; Sapindaceae)
-
Amazon Grape (Pourouma
cecropiaefolia; Moraceae)
-
Atemoya
-
Banana (Musacea
spp.; Musaceae); its starchy variant is the
plantain;
-
Barbados
Cherry (Malpighia glabra L.; Malpighiaceae),
also called Acerola, West Indian Cherry
-
Breadfruit (Artocarpus
altilis; Moraceae)
-
Burmese grape
(Baccaurea sapida; Euphorbiaceae)
-
Calabash (Crescentia
cujete; Cucurbitaceae)
-
CamuCamu (Myrciaria
dubia; Myrtaceae)
-
Carambola (Averrhoa
carambola; Oxalidaceae), also called star
fruit
-
Cempedak (Artocarpus
champeden; Moraceae)
-
Cherimoya (Annona
cherimola; Annonaceae)
-
Coconut (Cocos
spp.; Arecaceae)
-
Custard apple
(Annona reticulata; Annonaceae), also called
Bullock's Heart
-
Damson Plum (Chrysophyllum
oliviforme; Sapotaceae), also called Satin
Leaf
-
Date-plum (Diospyros
lotus)
-
Dragonfruit (Hylocereus
spp.; Cactaceae), also called pitaya
-
Durian (Durio
spp.; Bombacaceae)
-
Eggfruit (Pouteria
campechiana; Sapotaceae), also called
canistel or yellow sapote
-
Elephant apple
(Dillenia indica; Dilleniaceae)
-
Guarana (Paullinia
cupana; Sapindaceae)
-
Guavaberry or
Rumberry; (Myrciaria floribunda; Myrtaceae)
* Horned melon (Cucumis metuliferus;
Cucurbitaceae)
-
Indian Prune (Flacourtia
rukan; Flacourtiaceae)
-
Jackfruit (Artocarpus
heterophyllus Moraceae), also called nangka
-
Jambul (Syzygium
cumini)
-
Jambo?
(Eugenia malaccensis; Myrtaceae)
-
Jatobá (Hymenae
coubaril; ; Leguminosae Caesalpinioideae)
-
Jenipapo (Genipa
americana; Rubiaceae)
-
Keppel fruit (Stelechocarpus
burakol; Annonaceae)
-
Kandis (Garcinia
forbesii; Clusiaceae)
-
Kundong (Garcinia
sp.; Clusiaceae)
-
Langsat (Lansium
domesticum), also called longkong or duku
-
Mabolo, (Diospyros
discolor; Ebenaceae) also known as a velvet
persimmon
-
Mamey sapote (Pouteria
sapota; Sapotaceae)
-
Mamoncillo (Melicoccus
bijugatus; Sapindaceae), also known as
quenepa, genip or Fijian Longan
-
Mango (Mangifera
indica; Anacardiaceae)
-
Mangosteen (Garcinia
mangostana; Clusiaceae)
-
Marang (Artocarpus
odoratissima; Moraceae), a breadfruit
relative
-
Naranjilla,
Lulo (Solanum quitoense; Solanaceae)
-
Papaya (Carica
papaya; Caricaceae)
-
Peanut butter
fruit (Bunchosia argentea; Malpighiaceae)
-
Pequi or
Souari Nut (Caryocar brasiliense;
Caryocaceae)
-
Pineapple (Ananas
comosus or Ananas sativas; Bromeliaceae)
-
Pitomba (Talisia
esculenta ; Sapindaceae)
-
Pupunha or
peach-palm (Bactris gasipaes; Palmae)
-
Poha or Cape
Gooseberry (Physalis peruviana; Solanaceae)
-
Poisonleaf (Dichapetalum
cymosum) (?)
-
Rambutan (Nephelium
lappaceum; Sapindceae)
-
Red momblin (Spondias
purpurea; Anacardiaceae)
-
Riberry (Syzygium
luehmannii; Myrtaceae), also called Lilly
Pilly, Lillipilli, Chinese Apple
-
Rose apple
(Eugenia/Syzygium aquem; Myrtaceae), also
called Malay apple
-
Salak (Salacca
edulis), also called snakefruit
-
Sapodilla (Achras/Manilkara
zapota; Sapotaceae), also called chiku,
mespel, naseberry, sapadilla, snake fruit,
sawo
-
Soursop (Annona
muricata; Annonaceae), also called guanabana
-
Star apple (Chrysophyllum
cainito), also called caimito
-
Sugar apple (Annona
squamosa; Annonaceae)
-
Surinam Cherry
(Eugenia uniflora; Myrtaceae) also called
Brazilian Cherry, Cayenne Cherry, Pitanga
-
Tamarind (Tamarindus
indica; Caesalpiniaceae)
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